LONDON — There was three-time Grand Slam finalist and No. 3 seed Alexander Zverev, outplayed over five sets in a first-round loss at Wimbledon to 72nd-ranked Arthur Rinderknech, who entered Tuesday with a 1-4 career record at the All England Club and zero trips past the third round in 18 appearances at majors.

There was No. 7 Lorenzo Musetti, a semifinalist at Wimbledon last year and at the French Open last month, sent home Tuesday by Nikoloz Basilashvili, a qualifier ranked 126th who only once has made it as far as the fourth round in his 31 previous Grand Slam tournaments.

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And, most striking of all, as night arrived, there was No. 2 Coco Gauff going from the champion at Roland-Garros to a quick exit at Wimbledon, beaten 7-6 (3), 6-1 by Dayana Yastremska.

And on and on went the upsets on Day 2 at the grass-court major, meaning 23 seeds — 13 men, 10 women — failed to get to the second round, equaling the highest total at any Grand Slam tournament since they began assigning 32 seeds in each singles bracket in 2001.

US player Jessica Pegula reacts as she plays against Italy’s Elisabetta Cocciaretto during their women’s singles first round tennis match on the second day of the 2025 Wimbledon Championships at The All England Lawn Tennis and Croquet Club in Wimbledon, southwest London, on July 1, 2025. (Photo by Adrian Dennis / AFP) / RESTRICTED TO EDITORIAL USE

No. 3 Jessica Pegula also was among those leaving. The American was the runner-up at last year’s U.S. Open and was coming off a grass-court title in Germany over the weekend, defeating Iga Swiatek in the final, yet didn’t pose much of a challenge to 116th-ranked Elisabetta Cocciaretto in a 6-2, 6-3 loss that lasted less than an hour.

Two other major finalists, No. 5 Zheng Qinwen and No. 15 Karolina Muchova, were eliminated Tuesday, as were No. 26 Marta Kostyuk and No. 25 Magdalena Frech, whose opponent, 18-year-old Canadian Victoria Mboko, lost in qualifying last week and only got into the field when another player, Anastasia Potapova, withdrew with an injured hip.

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Nothing was quite as out-of-nowhere, though, as Rinderknech’s success. At least Yastremska has been a major semifinalist, at last year’s Australian Open.

“What a moment. Such emotions,” Rinderknech, a 29-year-old from France, said after completing his 7-6 (3), 6-7 (8), 6-3, 6-7 (5), 6-4 victory across 4 hours, 40 minutes against Zverev in a match suspended Monday night at a set apiece. “I don’t even know where to start.”

Coco Gauff of the U.S. reacts during her first round women’s single match against Dayana Yastremska of Ukraine at the Wimbledon Tennis Championships in London, Tuesday, July 1, 2025.(AP Photo/Kirsty Wigglesworth)

He ended things with a backhand winner, then dropped to his stomach, face down, on Centre Court.

Zverev joined Musetti — who hadn’t played since a leg injury forced him to stop at Roland-Garros and was a 6-2, 4-6, 7-5, 6-1 loser against Basilashvili — as top-10 losers on Tuesday, a day after No. 8 Holger Rune and No. 9 Daniil Medvedev departed.

Other seeded men exiting on Day 2 included No. 18 Ugo Humbert, No. 27 Denis Shapovalov, No. 28 Alexander Bublik and No. 30 Alex Michelsen.

Rinderknech pulled off his win thanks to some terrific serving, delivering 25 aces and saving all nine break points he faced. He converted three break chances against Zverev and won the point on 44 of his 55 trips to the net.

“It’s my first top-five win, in the biggest stadium in the world,” Rinderknech said. “My legs are still shaking. I’m just so happy the match is finished.”